Scripps Research’s Skaggs Graduate School awards doctoral degrees to 33rd graduating class

May 22, 2025


LA JOLLA, CA—Scripps Research’s Skaggs Graduate School of Chemical and Biological Sciences awarded doctoral degrees to 61 graduate students during the institute’s 33rd commencement on May 16, 2025. The ceremony represented the largest graduating class since the doctoral program’s inception in 1989.

“Today, we have the honor of celebrating 61 tremendously talented students who have completed their doctoral degrees,” said Keary Engle, PhD, dean of graduate and postdoctoral studies at Scripps Research, during the ceremony’s opening remarks. “Graduate students of this program have made vital contributions to science, from treatments for several types of cancer and increasing longevity to mapping how the human brain operates and the intricacies of chronic diseases. These students have expanded what we know and reimagined what is possible.”

This year’s class of PhD recipients also marked the 1,000th student to earn a doctoral degree since the program’s first commencement in 1993. Engle, himself a Scripps Research alumnus, hosted current and former graduates at a homecoming reception the day before commencement to celebrate the Class of 2025 and welcome them to the ranks of the institute’s alumni. At Friday’s ceremony, Engle spoke of their achievements and contributions to science during their time at Scripps Research.  

“We’ve already seen their impact resonate across the scientific community,” said Engle. “Their research has been cited over 9,000 times, and more than 220 journal articles list a student standing on the stage today as a contributing author. This cohort’s ability to translate sophisticated research ideas into successful funding proposals also was remarkable, with 23 competitive student fellowships awarded by organizations like the National Institutes of Health and the National Science Foundation.”

Members of this class also received support from 28 awards and endowed fellowships, which were created by Scripps Research board members and faculty, along with individuals and foundations that are passionate about scientific education. The number of endowed fellowships available for Scripps Research graduate students recently crossed the 100 mark, an ambitious milestone that began in 2018 with a lead gift from the Skaggs family and renaming the program the Skaggs Graduate School. Each endowed fellowship provides complete funding for one graduate student for the first three years of their studies, lifting much of the financial burden on their laboratory and enabling more of increasingly uncertain grant funding to be dedicated to critical research infrastructure and essential technologies. Institute leadership recently set a  goal of establishing endowments to provide this support for all Scripps Research graduate students in their first three years.

Engle also spoke about the unusual challenges faced by the graduating students, many of which embarked on graduate studies during or immediately before the COVID-19 pandemic.

“You’ve walked a path that was anything but predictable,” Engle told the graduates. “You experienced your graduate research and degree progression during a global pandemic. You adapted in the face of uncertainty, not just in your education, but to shifts in the world around you. That uncertainty continues today as we face unprecedented challenges to the future of science and reduction of funding. You're entering the world not only armed with knowledge, but with resilience, and that, perhaps more than anything, is what the current landscape of the world demands.”

Scripps Research President and CEO Pete Schultz, PhD, the L.S. “Sam” Skaggs Presidential Chair, echoed Engle’s sentiment with his remarks before the conferral of degrees.

“We really are in interesting times right now in science, and I think this class is better equipped than any class in the history of science with the tools and knowledge to have an impact not only on science but on society,” said Schultz. “I'd just like to remind the graduating class that it really is a privilege to do science but remember that you always have an obligation with a privilege, and that's ultimately to give back to those that are less fortunate than you, and I hope you take on that responsibility and burden.”

With Friday’s commencement, alumni of Scripps Research’s doctoral program increased to over 1,015. Many of this year’s graduates are extending their doctoral training into new research areas as postdoctoral scholars at universities and major academic research institutions. Others have joined companies in the pharmaceutical and bioscience industries, or start-ups focused on cutting-edge biotech research and drug discovery. Many students apply their knowledge and experience to different career paths, including food science, patent law and the venture capital and financial advising industries.

Profiles of many of this year’s graduates and a recording of the full ceremony can be viewed on the institute’s 2025 Graduation webpage, scripps.edu/graduation.

 

Skaggs Graduate School of Chemical and Biological Sciences
2025 Graduating Class

2025 Graduates

Thesis Advisors

Omar Apolinar

Keary M. Engle, PhD

Eleanor Bashian

James C. Paulson, PhD and Peng Wu, PhD

Nathan A. Beutler

Dennis R. Burton, PhD

Maya L. Bulos

Michael J. Bollong, PhD

Batuujin Burendei

Stefano Forli, PhD and Andrew B. Ward, PhD

Taiwei Chang

Jin-Quan Yu, PhD

Nikita Chekshin

Jin-Quan Yu, PhD

Kai-Yu Chen

Andrew I. Su, PhD

Wenqian Chen

Gabriel C. Lander, PhD

Megan L. Critcher

Mia L. Huang, PhD

Caroline A. Cuoco

Jeffery W. Kelly, PhD

Ngoc Thanh Nam (Nathan) Dao

Ryan A. Shenvi, PhD

Xiyu Dong

James W. Williamson, PhD

Ines Forrest

Christopher G. Parker, PhD

Brett Michael Garabedian

James C. Paulson, PhD

Quentin M. R. Gibaut

Susana T. Valente, PhD

Adriana Carolina Gonzalez-Cavazos

Andrew I. Su, PhD

Adrian Marco Guerrero

Benjamin F. Cravatt, PhD and Jeffery W. Kelly, PhD

Althea Theodora Danielle Hansel-Harris

Stefano Forli, PhD

Erika Marina James

Luke L. Lairson, PhD

Celina L Keating

Dennis R. Burton, PhD

Ahmed Khattab

Ali Torkamani, PhD and Eric P. Zorrilla, PhD

Mildred Apollo Kissai

Luke L. Lairson, PhD

Sergio Rodriguez Labra

Jeffery W. Kelly, PhD

Eleanor Marie Landwehr

Ryan A. Shenvi, PhD

André Nicolás León

Andrew B. Ward, PhD

Yihao Li

Jin-Quan Yu, PhD

Chung-Chih Liu

Supriya Srinivasan, PhD

Nicole Madrazo

R. Luke Wiseman, PhD

Colleen A. Maillie

Andrew B. Ward, PhD and Marco Mravic, PhD

Michaela Medina

Danielle A. Grotjahn, PhD

Rebecca M. Mello

Katja A. Lamia, PhD

Samantha M. Meyer

Matthew D. Disney, PhD

Karina Nugroho

Jeffery W. Kelly, PhD

Kayla Elaine Nutsch

Michael J. Bollong, PhD

Andi Pan

Michael Farzan, PhD and Mauricio A. Martins, PhD

Linghang Peng

David Nemazee, PhD

Michael L. Pigula

Peter G. Schultz, PhD

Nan Qiu

Alexander Adibekian, PhD

Brittney Racioppo

Alexander Adibekian, PhD

Camille Z. Rubel

Keary M. Engle, PhD

Roberto Modesto Salatino

Michalina Janiszewska, PhD

Philipp Nikolaus Sander

Luke L. Lairson, PhD

Yujie Shi

Peng Wu, PhD

Adam L. Sobel

Michael G. Constantinides, PhD

Carter N. Stout

Hans Renata, PhD

Amanda Sul

Luke L. Lairson, PhD

Mary Rita Tenuta

James E. Voss, PhD

Yuquan Tong

Matthew D. Disney, PhD

Shanel Mariko Tsuda

Matthew E. Pipkin, PhD

Ingrid Vallée

Xiang-Lei Yang, PhD

Elliot Allen Warner

Peter G. Schultz, PhD

Christopher Conrad Williams

Douglas J. Kojetin, PhD

Adrianna Nicole Wilson

Laura A. Solt, PhD

Yin Wu

Sandra E. Encalada, PhD

Tong Wu

Xiaohua Wu, PhD

Yi Xie

Hollis T. Cline, PhD and Anton Maximov, PhD

Shaochen You

Michael J. Bollong, PhD

Jinhan Yu

Donna G. Blackmond, PhD

Zixuan Yuan

Scott B. Hansen, PhD and Sathyanarayanan V. Puthanveettil, PhD

Yuanjin Zhang

Benjamin F. Cravatt, PhD

About the Skaggs Graduate School

Ranked among the top 10 doctoral programs of its kind in the nation by U.S. News & World Report, the Skaggs Graduate School of Chemical and Biological Sciences at Scripps Research offers rigorous training in chemistry, chemical biology, neuroscience, immunology, computational biology, and numerous other biomedical research areas. The program immerses students in intensive laboratory research while offering a customizable course curriculum that allows students to match individual research interests while exploring multidisciplinary topics at the interface of chemistry and biology. 


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